Ionis Pharmaceuticals Inc (IONS)

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Institutional Investors' Holdings

Filing Period: Filing is Ongoing

No.	Name	Shares	Value	% Port
1.	Fmr Llc	24,740,108	$1,104,645,822	0.05%
2.	AmundiBOND	3,450,000	$377,640,391	0.08%
3.	Geode Capital Management, Llc	3,277,569	$146,379,160	0.01%
4.	Pictet Asset Management Holding Sa	1,944,020	$87,249,218	0.08%
5.	Tweedy Browne Co Llc	1,793,400	$80,075,310	6.52%