Ionis Pharmaceuticals Inc (IONS)

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Institutional Investors' Holdings

Filing Period:

No.	Name	Shares	Value	% Port
1.	Fmr Llc	23,613,585	$712,421,869	0.05%
2.	Vanguard Group Inc	16,383,934	$494,303,288	0.01%
3.	Capital World Investors	12,361,994	$372,961,359	0.06%
4.	Blackrock, Inc.	11,264,442	$339,848,216	0.01%
5.	Wellington Management Group Llp	8,416,421	$253,923,422	0.05%